rs121434467

Variant summary

Our verdict is Uncertain significance. The variant received 4 ACMG points: 4P and 0B. PS4_ModeratePS3_Moderate

This summary comes from the ClinGen Evidence Repository: The m.4295A>G variant in MT-TI has been reported in at least four families from different ethnic groups with features of primary mitochondrial disease including deafness, hearing loss, and cardiomyopathy (PMIDs: 18177739, 22241583, 33398350, 34991096, 8889580; PS4_moderate). Of note, other isolated phenotypes have been reported including essential hypertension and autism spectrum disorder, but these are outside the scope of this curation (PMIDs: 11406419, 19778529, 19043581). This variant is seen in the general population (<0.5% in three large population databases: Mitomap - 112/61168, gnomAD v3.1.2 - 167/56431, and Helix - 923/195983). Notably, these occurrences are almost exclusively in haplogroup (hg) K (110/112 in specifically K1a in Mitomap; 166/167 in hg K in gnomAD, 911/923 in K in Helix) and these three databases combined reported only 15 individuals from non-K haplogroups (H, I, R, X, J, U, and V). In silico predictors are conflicting as the computational predictor MitoTIP suggests this variant is possibly benign (0.44) but HmtVAR predicts it to be pathogenic (0.75). Of note, this variant is 100% conserved among 45 species ranging from Drosophila to Homo sapiens in the extraordinarily conserved anticodon loop of tRNA Isoleucine. Cybrid studies showed multiple deleterious biochemical impacts of the variant in a Chinese family from haplogroup D4j (PMID:33398350). Other studies showed this variant reduces the efficiency with which tRNAIle can be processed by 3′-tRNase (PMID:18177739). An in-vitro system utilizing plasmids with tRNAIle inserts showed that this variant reduces the tRNase Z reaction 10.3-fold (PMID:12655007). On the basis of these strong, independent, and consistent functional studies, this Expert Panel elected to give higher weight to this line of evidence (PS3_moderate). In summary, this variant meets criteria to be classified as uncertain significance for primary mitochondrial disease inherited in a mitochondrial manner. We note there was much discussion of the possible differing roles of this variant in haplogroups K1a (protective or tolerated) and D4j (damaging), however the VCEP elected to keep the classification of this variant as uncertain significance (four of 11 experts felt likely benign was the more appropriate classification). This classification was approved by the NICHD/NINDS U24 ClinGen Mitochondrial Disease Variant Curation Expert Panel on December 11, 2023. Mitochondrial DNA-specific ACMG/AMP criteria applied (PMID:32906214): PS4_moderate, PS3_moderate. LINK:https://erepo.genome.network/evrepo/ui/classification/CA254841/MONDO:0044970/015

Frequency

Mitomap GenBank:
𝑓 0.0018 ( AC: 113 )

Consequence

TRNI
unassigned_transcript_4790 synonymous

Scores

Mitotip
Uncertain
12

Clinical Significance

Uncertain significance reviewed by expert panel P:2U:5B:1
MHCM-/-Maternally-inherited-hypertension-/-Maternally-inherited-deafness

Conservation

PhyloP100: 3.44

Publications

7 publications found
Variant links:
Genes affected
TRNI (HGNC:7488): (mitochondrially encoded tRNA isoleucine)
MT-ND2 (HGNC:7456): (mitochondrially encoded NADH dehydrogenase 2) Enables NADH dehydrogenase (ubiquinone) activity. Involved in mitochondrial electron transport, NADH to ubiquinone and mitochondrial respiratory chain complex I assembly. Part of mitochondrial respiratory chain complex I. Implicated in Leber hereditary optic neuropathy; multiple sclerosis; myocardial infarction; neurodegenerative disease (multiple); and urinary bladder cancer. [provided by Alliance of Genome Resources, Apr 2022]
MT-ND1 (HGNC:7455): (mitochondrially encoded NADH dehydrogenase 1) Enables NADH dehydrogenase (ubiquinone) activity. Involved in mitochondrial electron transport, NADH to ubiquinone and mitochondrial respiratory chain complex I assembly. Located in mitochondrial membrane. Part of mitochondrial respiratory chain complex I. Implicated in several diseases, including MELAS syndrome; neurodegenerative disease (multiple); optic nerve disease (multiple); toxic shock syndrome; and type 2 diabetes mellitus. Biomarker of Alzheimer's disease; Parkinson's disease; and multiple sclerosis. [provided by Alliance of Genome Resources, Apr 2022]
TRNM (HGNC:7492): (mitochondrially encoded tRNA methionine)
TRNQ (HGNC:7495): (mitochondrially encoded tRNA glutamine)
TRNQ Gene-Disease associations (from GenCC):
  • mitochondrial disease
    Inheritance: Mitochondrial Classification: LIMITED Submitted by: ClinGen

Genome browser will be placed here

ACMG classification

Classification was made for transcript

Our verdict: Uncertain_significance. The variant received 4 ACMG points.

PS3
For more information check the summary or visit ClinGen Evidence Repository.
PS4
For more information check the summary or visit ClinGen Evidence Repository.

Variant Effect in Transcripts

ACMG analysis was done for transcript: ENST00000387365.1. You can select a different transcript below to see updated ACMG assignments.

RefSeq Transcripts

Selected
GeneTranscriptTagsHGVScHGVSpEffectExon RankProteinUniProt

There are no transcript annotations for this variant.

Ensembl Transcripts

Selected
GeneTranscriptTagsHGVScHGVSpEffectExon RankProteinUniProt
MT-TI
ENST00000387365.1
TSL:6
n.33A>G
non_coding_transcript_exon
Exon 1 of 1
MT-ND2
ENST00000361453.3
TSL:6
c.-175A>G
upstream_gene
N/AENSP00000355046.4
MT-ND1
ENST00000361390.2
TSL:6
c.*33A>G
downstream_gene
N/AENSP00000354687.2

Frequencies

Mitomap GenBank
AF:
0.0018
AC:
113
Gnomad homoplasmic
AF:
0.0030
AC:
167
AN:
56431
Gnomad heteroplasmic
AF:
0.0
AC:
0
AN:
56431
Alfa
AF:
0.00201
Hom.:
46

Mitomap

Disease(s): MHCM-/-Maternally-inherited-hypertension-/-Maternally-inherited-deafness
Status: Reported-[VUS]
Publication(s): 8889580

ClinVar

ClinVar submissions as Germline

Significance:Uncertain significance
Revision:reviewed by expert panel
View on ClinVar
Pathogenic
VUS
Benign
Condition
-
3
-
not provided (3)
-
-
1
MELAS syndrome (1)
-
1
-
Mitochondrial disease (1)
1
-
-
Mitochondrial non-syndromic sensorineural hearing loss (1)
-
1
-
not specified (1)
1
-
-
Primary familial hypertrophic cardiomyopathy (1)

Computational scores

Source: dbNSFP v4.9

Name
Calibrated prediction
Score
Prediction
Mitotip
Uncertain
12
Hmtvar
Pathogenic
0.75
PhyloP100
3.4

Publications

Other links and lift over

dbSNP: rs121434467; hg19: chrM-4296; API